[19] Grobis, M. M., Jolles, J. W., Mazué, G. P. F., Twomey, C. R., Bak-Coleman, J., Rubenstein, D. I., Couzin, I. D. (In prep) Quantifying the spatial distribution of risk in fish schools

[18] Jolles, J. W., & Killen, S. (In prep) Individual heterogeneity in animal collectives: Mechanisms and consequences across social scales

[17] Jolles, J. W., Briggs, H., Araya-Ajoy, Y., Manica, A., Boogert, N. J. (In prep) Personality, plasticity and predictability in sticklebacks: bold fish are less plastic and more predictable than shy fish.


[16] Jolles, J. W. (2018) Comment on ‘Consistency of fish‐shoal social network structure under laboratory conditions (Gaffney & Webster, 2018)’. Journal of Fish Biology [link]

[15] Jolles, J. W., Laskowski, K, Boogert, N. J., Manica, A. (2018) Repeatable group differences in the collective behaviour of stickleback shoals across ecological contexts. Proceedings of the Royal Society B 285: 20172629. Doi: 10.1098/rspb.2017.2629 [linkopen access

[14] Daversa, D. D., Manica, A., Bosch, J., Jolles, J. W., Garner, T. W. J. (2018) Routine habitat switching alters the likelihood and persistence of infection with a pathogenic parasite. Functional Ecology [link]


[13] Jolles, J. W., Boogert, N. J., Sridhar, V., Couzin, I. D., Manica, A. (2017) Consistent individual differences drive collective behaviour and group functioning of schooling fish. Current Biology 27: 1-7. [pdfopen access.
Find a nice Dispatch article on our paper in Current Biology by James Herbert-read.


[12] Greggor, A. L., Jolles, J. W., Thornton, A., Clayton, N. S. (2016) Seasonal changes in neophobia and its consistency in rooks: the effect of novelty type and dominance position. Animal Behaviour 121: 11:20 [pdf]

[11] Jolles, J. W., Manica, A., Boogert, N. J. (2016) Food intake rates of inactive fish are positively linked to boldness in three-spined sticklebacks. Journal of Fish Biology. 88(4): 1661-1668 [pdfopen access

[10] Jolles, J. W., Aaron Taylor, B., Manica, A. (2016) Recent social conditions affect boldness repeatability in individual sticklebacks. Animal Behaviour 112: 139-145 [pdf]


[9] Jolles, J. W., Boogert, N, van den Bos, R. (2015). Sex differences in risk-taking and associative learning in rats. Royal Society Open Science 2: 150485. [pdf]

[8] Jolles, J. W., Wilson, A., Stumpe, M. C., Johnstone, R.A., Manica, A. (2015). The role of social attraction and its link with boldness in the collective movements of three-spined sticklebacks. Animal Behaviour 99: 147-153. pdf] [BBC Radio5Live at 01:26:01] [See a 4min video about our paper here]


[7] Jolles, J. W., Wilson, A., Stumpe, M. C., Johnstone, R.A., Manica, A. (2014). The role of previous social experience on risk-taking and leadership in three-spined sticklebacks. Behavioral Ecology 25(6): 1395-1401. [pdf]

[6] Zandberg, L., Jolles, J. W., Boogert, N. J., Thornton., A. (2014). Jackdaw nestlings can discriminate between conspecific calls but do not beg specifically to their parents. Behavioral Ecology. 25(3): 565–573 [pdf]


[5] van den Bos, R., Jolles, J. W. & Homberg, J. (2013) Social modulation of decision-making: a cross-species review. Frontiers in Neuroscience 7(301). [pdf]

[4] Jolles, J. W., Ostojic, L & Clayton, N. S. (2013) Dominance, pair bonds and boldness determine social foraging tactics in rooks (Corvus frugilegus). Animal Behaviour. 85(6): 1261-1269. [pdf]

[3] Jolles, J. W., King, A.J., Manica, A., Thornton, A. (2013) Heterogeneous structure in mixed-species corvid flocks in flight. Animal Behaviour 85(4): 743-750. [pdf] [Planet Earth] [BirdGuides]


[2] van den Bos, R., Jolles, J. W., van der Knaap, L., Baars, A., de Visser, L. (2012) Male and female Wistar rats differ in decision-making performance in a rodent version of the Iowa Gambling Task. Behavioural Brain Research 234(2): 375-379. pdf]


[1] Jolles, J. W., de Visser, L., van den Bos, R. (2011) Male Wistar rats show individual differences in an animal model of Conformity. Animal Cognition 14: 769-773. [pdf]